2009 - | Emeritus Professor | Department of Biology, University of York |
1989 - | Professor | Department of Biology, University of York |
1980 - 1989 | Reader | Department of Biology, University of York |
1973 - 1980 | Senior Lecturer | Department of Biology, University of York |
1965 - 1973 | Lecturer | Department of Biology, University of York |
1965 | PhD | Imperial College, London University |
1964 | Diploma in Biochemistry | Chelsea College, London University |
1962 | BSc Zoology | Imperial College, London University |