|
2009-Present |
Honorary Fellow |
Department of Biology, University of York |
|
1972-2009 |
Lecturer/Senior Lecturer in Population Genetics |
Department of Biology, University of York |
|
1971-1972 |
Research Demonstrator |
Department of Genetics, University of Swansea |
|
1968-1971 |
PhD |
Department of Genetics, University of Liverpool |
|
1965-1968 |
BSc |
Department of Zoology, University College of North Wales, Bangor |
My research has focussed on three separate themes: (1) the maintenance and evolutionary significance of colour polymorphisms in spiders; (2) speciation and hybridisation in spiders; (3) ecology and conservation of the endangered Tansy beetle.

Some colour morphs of the Hawaiian happy-face Theridion grallator
