Dr Pen Holland
Lecturer in Ecology



2016 - present Lecturer Univeristy of York
2015 - 2016 Associate Lecturer University of York
2014 - 2015 Teaching Fellow in Ecology University of York
2010 - 2014 Research Scientist     Landcare Research, New Zealand
2009 - 2010 Postdoctoral Researcher Landcare Research, New Zealand
2005 - 2008 PhD (Biology) University of York
2003 - 2004 MRes (Mathematics in the Living Environment)   University of York
1999 - 2003 BSc (Mathematics) University of York

Holland Pen

Contact details

Dr Pen Holland
Lecturer in Ecology
Department of Biology
University of York
YO10 5DD

Tel: 01904 328734



Key themes running through my research are: finding simple models to understand complex ecological problems; using noisy or sparse data to find quantitative solutions for real life problems; and developing novel ways to teach, disseminate research, and engage people in the biosciences. 
Two broad areas of research are currently of interest to me:
  • Quantifying and predicting in ecology: spatial patterns, resources, dispersal and connectivity across landscapes, probability of detection issues, and effective use of scarce, uncertain or ordinal data. Real world applications include models to assess and predict herbivore density-damage impacts on plants, and interactions between climate, plant productivity and animal populations in resource pulse-driven ecosystems.
  • Games with a purpose: multiplayer games to understand perceptions of, and educate about, sustainable and economic use of shared resources, and as a communication conduit between multiple stakeholders in wicked ecological problems, and games to motivate, support and improve learning in higher education.